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The DEFCON level is controlled primarily by the U.S. President and the U.S. Secretary of Defense through the Chairman of the Joint Chiefs of Staff and the Combatant Commanders, and each DEFCON level defines specific security, activation and response scenarios for the troops in question.
Notably, it allows the combatant commander to more rapidly establish joint task forces and to better understand the battlespace before he uses his forces, thus giving him an advantage of time, perhaps his most critical resource.