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The fossil record of this family extends back to the Early Carboniferous.
The holotype was collected in the East Kirkton Quarry, near Bathgate of West Lothian, from "Bed 82" East Kirkton Limestone of the Bathgate Hills Volcanic Formation, dating to the Brigantian substage, of the late Viséan stage, of the Dinantian series (Early Carboniferous), about 333-328.3 million years ago.
The Widmerpool Gulf is a name given to a sedimentary basin that existed as an area of open water during the Early Carboniferous (Tournaisian Age).