In 1986, Cai graduated from the Department of Mathematics, Hangzhou University (previous and current Zhejiang University) when he was only 18 years old.
He holds a BS in biochemistry and microbiology from Hangzhou University in China; a PhD in biochemistry and a postdoctorate training in human genetics from Boston University; and an MS in software engineering from Brandeis University.
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In 1982, Guan graduated (B.S.) from the Department of Biology, Hangzhou University (previous and current Zhejiang University).