Llywelyn the Last | Gruffydd ap Llywelyn | Llywelyn the Great | Llywelyn | Iorwerth Peate | Iorwerth | Morgan Llywelyn | Madog ap Llywelyn | Llywelyn the Elder ap Maredudd ap Cynan ab Owain Gwynedd | Llywelyn Siôn | Llywelyn Bren | Llywelyn ap Dafydd | Iorwerth Drwyndwn |
She was the daughter of Domhnall I, Earl of Mar and Helen (or Ellen) of Wales (1246–1295), the illegitimate daughter of Llywelyn ab Iorwerth ("the Great") Prince of Wales; she had previously been the wife of Maol Choluim II, Earl of Fife.
The de Boulers (Bowdler) family held the castle until 1215 when the fortress was destroyed by Prince Llywelyn ab Iorwerth.