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The cars were built for services in the LMR Central Division and in the Liverpool - St Helens area, where the gradients in the Lancashire & Yorkshire area required more power.
The Region consisted principally of ex-Great Western Railway lines, minus certain lines west of Birmingham, which were transferred to the London Midland Region in 1963 and with the addition of all former Southern Railway routes west of Exeter, which were subsequently rationalised.
In a major national boundary change in 1958 the former Great Central network except those lines in Yorkshire and Lincolnshire transferred to the London Midland Region; in return the Eastern gained the former LMS lines in Lincolnshire and the present-day South Yorkshire.