The natural final host is the rat (Rattus rattus) and the intermediate host is the pomatiopsid snail Tricula bollingi.
He also performed Worth the Dog in the Woolworths adverts (again opposite Don Austen as Wooly The Sheep), Mervin J Minky on MTV's Fur TV and Rattus Rattus on the CBBC Horrible Histories series.
Serological studies have revealed, but not confirmed, other rodents in Thailand as possible reservoirs, including the black rat (Rattus rattus), Polynesian rat (R. exulans), brown rat (R. norvegicus) and lesser rice-field rat (R. losea).