Richard Goldschmidt in 1929 due to his experiments on drosophila through exposure to elevated temperatures had claimed to have produced evidence for directed mutation.
Davidse and MartÃnez suggested that L. schismatica could be one of Richard Goldschmidt's "hopeful monsters", meaning that the inverted floral morphology could have arisen from a macromutation in the genes that control floral development.
With the help of Richard Goldschmidt, she overcame gender barriers and started running her own laboratory in Berlin.
US geneticist Richard Goldschmidt wrote about him: "There was the brilliant Nikolai Koltsov, probably the best Russian zoologist of the last generation, an enviable, unbelievably cultured, clear-thinking scholar, admired by everybody who knew him".
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