Extensive long-term studies have been carried out on its foraging behaviour, in northern Germany, in the Netherlands and particularly on the River Exe estuary in south-west England.
Maintaining a studio in Exeter, he produced works around the Exe estuary, before being offered the post of Associate Artist at the National Gallery.
The Exe was named after the River Exe, although Rolls-Royce later transferred the using of river names to its gas turbine engines.
He was the Rector of the Exe Valley Group of Churches from then until 1972; and after that of Shirwell with Loxhore.
Isambard Kingdom Brunel, in surveying the South Devon Railway, opted to push a line along a coastal strip between the Exe and Teign valleys, and then to climb the southern outliers of Dartmoor making for the head of the Plym estuary.
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This also applies to the similarly spelled rivers Axe, Exe and Usk with the changes to the name evolving over ten centuries of usage.
Through the square runs a small stream which is one of many local tributaries of the River Clyst, which in turn feeds into the Exe.