For example, Darden examines the Mendelian "Theory of the Gene," which underwent significant theoretical modifications from around 1900 (when Mendel was "rediscovered") to 1926 (when Thomas Hunt Morgan published his famous textbook defining Mendelian theory in its roughly modern form).
In 1911 the American geneticist Thomas Hunt Morgan observed crossover in Drosophila melanogaster meiosis and provided the first genetic evidence that genes are transmitted on chromosomes.
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This view was expressed in the writings of key founders of genetics, including Thomas Hunt Morgan, Reginald Punnett, Wilhelm Johannsen, Hugo de Vries, William Bateson and others.