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He was the eldest son of son of Henry Colley and Mary, daughter of Sir William Ussher; brother of Henry Colley, father of Garret Wesley, 1st Earl of Mornington, father-in-law to Chichester Fortescue and William Francis Crosbie, and grandfather of Arthur Wellesley, 1st Duke of Wellington and Richard Wellesley, 1st Marquess Wellesley.