It is found in Agaricomycetes fungi in the orders Agaricales and Thelephorales.
He became a leading specialist in various areas, in which he enjoyed close collaborations: Agaricomycetes, with Quélet and Adalbert Ricken; Aphyllophoromycetideae, with Narcisse Théophile Patouillard and the Canon Hubert Bourdot; and finally Discomycetes with Boudier.