G. Evelyn Hutchison, studied the way logistic growth, biological feedback systems and self-regulation tended to explain many of the features of ecological systems, and Raymond Lindeman has further extended the way energy flows between various trophic levels in his "trophic-dynamic" model, further developed by Mark McMenamin and Dianna McMenamin's thesis of "Hypersea", which looks at the rate of water flow through the Gaian biological environment.