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The first elements of the 494th arrived on Angaur on 16 October, being strafed by Japanese forces still on the island as the battle of Bloody-Nose Ridge was still ongoing.
Haldane was shot by a Japanese sniper on October 12, 1944, while assessing the area of Hill 140 during the Battle of Peleliu in the Palau Islands, three days before the Marines came off the lines.
In the meantime, in order to secure the flank for U.S. forces preparing to attack Japanese forces in the Philippines, in September, 1944, U.S. Marine and Army forces landed on the islands of Peleliu and Angaur in Palau.