Stillman was educated at Glen Waverley High School (1966-1969) and Sydney Boys High School (1970–71), then graduated with First Class honours from the University of Sydney, and earned his Ph.D. from the John Curtin School of Medical Research at the Australian National University.
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He developed a biochemical system to study DNA replication-coupled chromatin assembly in a test tube and discovered proteins such as Chromatin Assembly Factor-1 (CAF-1) that cooperate with the DNA replication machinery to assemble new histones onto the DNA.
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