On the genus Carabus, he published a Catalogue of the types of the Paris Museum collection.
Carabus hummeli middendorfi is a subspecies of ground beetle in the Nebriinae subfamily that is endemic to Russia, where it can only be found on Bolshoy Shantar Island in Primorsky Krai.
Carabus nemoralis is a beneficial predator as it eats the agricultural pest Deroceras reticulatum/gray garden/milky slug, in its young stage and also its eggs.
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Carabus nemoralis is a ground beetle common in central and northern Europe, as well as Iceland and the island of Newfoundland.
Carabus obsoletus uhligi is a subspecies of ground beetle in the Carabinae subfamily that can be found in Hungary and Romania.