with Axel Leonard Melander (1878-1962), A Key to the Families of North American Insects (1915).
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He was appointed field agent of the Bureau of Entomology, United States Department of Agriculture 1904-05, curator of invertebrate zoology in the Milwaukee Public Museum 1905-09, and then became instructor in economic entomology at Harvard University.
Wheeler began studying ants at the Barro Colorado Island research station in the Panama Canal in 1924, having been urged to go to the tropics by William Morton Wheeler, who had mentored him at Harvard together with Charles Thomas Brues.
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