He had taken the doctorate in 1935 with the thesis Lebenszyklus, Cytologie und Sexualität der Chiorophycee Cladophora Suhriana Kiltzing, on the algae Cladophora.
Luang Prabang kháy sheets are the most readily available form of Mekong Weed and are famous throughout the country and in the neighbouring Isaan, though difficult to find beyond Vientiane.
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Quagga mussel populations have also increased tremendously in the same time frame.