A similar phenomenon is found in some butterflies (for example, several species of Colias and Gonepteryx, both of the family Pieridae).
Colias | Colias sifanica | Colias leechi | Colias erate | Colias caucasica |
Colias balcanica is often listed as a subspecies of Colias caucasica.
Colias nebulosa is much darker than Colias sifanica ; the forewing without middle spot, but with light spot at the distal margin; the hindwing almost uniformly black, with large,conspicuous, light yellow middle spot, and light ray-like spots on the veins.
Colias staudingeri leechi Grum-Grshimailo, 1893 (Ladakh) uncertain taxon
There is some dispute whether it ought to be included in the Eastern Pale Clouded Yellow (C. erate) as a subspecies Colias erate nilagiriensis, or treated as a distinct species Colias nilagiriensis.