In 1907 he was the first scientist to show that the complement could be split into two components that were singularly inactive, only regaining their activity when reunited.
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Membrane Inhibitor of Reactive Lysis, CD59, a cell surface glycoprotein that inhibits complement-mediated lysis.
In the circulation, Protein S exists in two forms: a free form and a complex form bound to complement protein C4b-binding protein (C4BP).