Livingston was born in Waycross, Georgia, and received a Bachelor of Arts, magna cum laude, from the Woodrow Wilson School of Public and International Affairs at Princeton University in 1980 and a Juris doctor from Harvard Law School in 1984, where she served as an editor of the Harvard Law Review.
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From 1986 to 1991, she was an assistant U.S. attorney in the Southern District of New York, where she handled criminal cases, including the prosecution of Ferdinand Marcos, former president of the Philippines.
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