It is often associated with mosses such as Brachythecium albicans, B. mutabulum, Campylopus introflexus, Ceratodon purpureus, Dicranum scoparium, Eurhynchium hians, E. praelongum, E. speciosum, Rhacomitrium canescens, Pohlia species or Polytrichum piliferum.
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