There is a single foramen on the dorsal side of the tarsometatarsus, with a plantar exit hole between the third and fourth metacarpal's distal ends (presumably for the outer toe's adductor tendon) and another (presumably for nerves and blood vessels) on the plantar surface of the tarsometatarsus.
•
The reed grasses of the subfamily Arundinoideae were probably far less widespread in Africa 35 million years ago (Ma) than they are today, but it is fairly likely that other Poales reeds—Cyperaceae (maybe including the ancestors of the Papyrus Sedge Cyperus papyrus), Juncaceae and Typhaceae—grew in aquatic habitats back then already.