These expeditions have taken him across many regions of the circumpolar Arctic, including the Northwest Territories and Nunavut in Canada; Fennoscandia; Greenland; Chukotka in Russia; and the Arctic National Wildlife Refuge in Alaska, USA.
It is found from Fennoscandia to the Kamchatka Peninsula and in northern North America, where it occurs across the boreal forest region, from Alaska across the Northwest Territories and Nunavut to Newfoundland, and in the mountains south to southern Wisconsin, Alberta and British Columbia.