Alexander von Humboldt said about howler monkeys, "their eyes, voice, and gait are indicative of melancholy", while John Lloyd Stephens described those at the Maya ruins of Copán as "grave and solemn, almost emotionally wounded, as if officiating as the guardians of consecrated ground".
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The hyoid of Alouatta is pneumatized, one of the few cases of postcranial pneumaticity outside Saurischia.
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