These prayers were written for a female reader, but the singular feminine Latin endings were first overwritten with singular masculine endings, and then still later, plural masculine ones.
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For most of its history, it was thought to have been the product of a scriptorium in the north of England, owing to its inclusion of a number of northern saints such as Oswald of Northumbria and John of Beverley (who very seldom occur outside northern manuscripts), although modern scholarly consensus puts its likely origin in the southwest of England.
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