Both Todd and Dutton caught the disease, but were well enough to continue traveling, and reached Kasongo on 9 February 1905.
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He died in the Congo Free State at the age of 29 from tick fever, or African relapsing fever, while investigating the disease, which is caused by a spirillum that was later named Borrelia duttoni.
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Dutton found that the monkeys could be infected by bites from soft ticks (Ornithodoros moubata) carrying Borrelia duttoni, a spirochaete.
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