Mike Cullen began his academic career by studying mathematics at Wadham College, Oxford, but later switched to zoology, spending time at the Edward Grey Institute of Field Ornithology while investigating the ecology of Marsh Tits.
marsh | Rod Marsh | Jordan Marsh | Stan Marsh | Marsh Harrier | Marsh Fritillary | Garry Marsh | Shaun Marsh | Othniel Charles Marsh | Moreton-in-Marsh | Bacchus Marsh | Romney Marsh | Ngaio Marsh | Natasha Marsh | Marsh & McLennan Companies | Marsh Creek State Park | Geoff Marsh | Dave Marsh | Tit for tat | Tawny Tit-Spinetail | Richard Marsh, Baron Marsh | Richard Marsh | P'tit quinquin | Mitchell Marsh | Michelle Marsh | Michael Marsh | Marsh Lake | Marsh | Mae Marsh | James Holt Marsh |
This is supported by mtDNA cytochrome b sequence analysis; within Poecile, the Willow Tit is part of a group of Eurasian "typical chickadees" that includes also the Marsh Tit, Père David's Tit, and the Songar Tit (Gill et al. 2005, del Hoyo et al. 2007).