Born prematurely in a suburb of Denver, Colorado, Mary MacPherran grew up scrawny and short-statured, inspiring a popular girl at school, Vanessa Ashwood, to nickname her "Skeeter" (a slang word for mosquito).
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Battered and inadvertently humiliated by her, Titania was offered the chance at revenge after obtaining the Power Gem, one of the legendary Infinity Gems, from its former owner, the former Champion of the Universe, now known as the Fallen One because of his own defeat at the She-Hulk's hands.
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However, this stay in The Vault was as permanent as earlier ones – a conspiracy between Loki and the Wizard saw Titania free once more, in Loki's Acts of Vengeance plan.
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