The original mutations in the ptc gene were discovered in the fruit fly Drosophila melanogaster by 1995 Nobel Laureates Eric F. Wieschaus and Christiane Nusslein-Volhard and colleagues, and the gene was independently cloned in 1989 by Joan Hooper in the laboratory of Matthew P. Scott, and by Philip Ingham and colleagues.
He returned to the UK in 1982, joining the Imperial Cancer Research Fund (ICRF, now known as Cancer Research UK) as a post-doctoral research fellow in the laboratory of David Ish-Horowicz.
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After a brief spell as a Research Scientist at the MRC Laboratory of Molecular Biology in Cambridge, he re-joined the ICRF as a staff scientist, remaining there for ten years before moving to the University of Sheffield, where he established the MRC Centre for Developmental and Biomedical Genetics.
The founding member of TrxG Proteins, trithorax (trx), was discovered by P.W. Ingham as part of his doctoral thesis while a graduate student in the laboratory of J.R.S. Whittle at the University of Sussex.
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