In the early 1990s, Karlin and Stephen Altschul developed the Karlin-Altschul statistics, a basis for the highly used sequence similarity software program BLAST.
Most notably, Altschul is the co-author of the BLAST algorithm used for sequence analysis of proteins and nucleotides.
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He is the son of Stephanie Rosemary (née Wagner) and Arthur Altschul, a former partner at Goldman Sachs.
The empirical use of Sum statistics in the treatment of gapped alignments was validated in collaboration with Stephen Altschul, from 1994-1995.
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