From 1925 to 1950 he was based at the Carnegie Institute of Washington, at Stanford University where he worked on plant species concepts with Jens Clausen and William Hiesey.
With taxonomist David D. Keck and physiologist William Hiesey, he formed the first interdisciplinary effort to combine genetics, ecology and systematics in order to understand the ecological genetics of the evolutionary process in California plants.
William Shakespeare | William Laud | William Blake | William | William III of England | William Morris | William McKinley | William Howard Taft | William Ewart Gladstone | William the Conqueror | William S. Burroughs | William Shatner | William Faulkner | William Randolph Hearst | William Wordsworth | William Tecumseh Sherman | William Hogarth | Prince William, Duke of Cambridge | William Penn | William Jennings Bryan | William Gibson | William Wilberforce | William James | William Makepeace Thackeray | Fort William | William Hanna | William Hague | William III | William Hurt | William Walton |