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William won his patrimony in a war with his half-brother Odo, who was killed in battle at Mauzé.
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Two daughters have been hypothesized to be children of this couple: Clementia, who married Conrad I of Luxembourg, and Agnes, who married Peter I of Savoy.
Having become the military leader of various Lombard cities, including Pavia, Vercelli, Alessandria, Tortona, Genoa, Turin, Asti, Alba, Novara, Brescia, Cremona, and Lodi, he was also elected head of the anti-Angevin coalition.