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The most damage however was caused by habitat destruction and the other animals humans brought with them, particularly rats (the Polynesian rat or kiore introduced by Māori and the Brown Rat and Black Rat subsequently introduced by Europeans), but also mice, dogs, cats, stoats, weasels, pigs, goats, deer, hedgehogs, and Australian possums.
Serological studies have revealed, but not confirmed, other rodents in Thailand as possible reservoirs, including the black rat (Rattus rattus), Polynesian rat (R. exulans), brown rat (R. norvegicus) and lesser rice-field rat (R. losea).