Besides special articles on the morphology and cytology of algæ, fungi, and liverworts, and studies in the Œnothera, he was coauthor with J. Y. Bergen of Principles of Botany (1906) and Laboratory and Field Manual of Botany (1907).
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He held a position at the Marine Biological Laboratory, Woods Hole, Mass. (1897–1905) and at the Bureau of Fisheries.
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