The British War Office showed no interest until 1940, when a prototype was constructed using a Matilda II tank.
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They landed in Normandy on 12 August 1944, and saw no action until 29 September 1944 when they were ordered to transfer all their equipment to the 42nd and 49th Royal Tank Regiments, and were retrained to operate the American amphibious LVT-4, known by the British Army as the Buffalo Mark IV.
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The unit trained at Lowther Castle near Penrith, and were based at Brougham Hall, Cumberland, spent 1942 and 1943 in the Middle East without seeing action, and returned to the UK in April 1944.
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