He invented three methods for the estimation of variance components in unbalanced settings of mixed models, and invented a method for constructing the inverse of Wright's numerator relationship matrix based on a simple list of pedigree information.
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He, with his Ph.D. student Shayle R. Searle, greatly extended the use of matrix notation in statistics.
Charles Darwin | Charles Dickens | Charles, Prince of Wales | Ray Charles | Charles II of England | Charles I of England | Charles Lindbergh | Charles de Gaulle | Charles II | Charles | Charles I | Roy Rogers | Prince Charles | Charles V | Roy Orbison | Joe Henderson | Charles Scribner's Sons | Charles Aznavour | Roy Lichtenstein | Charles University in Prague | Charles Stanley | Charles Bukowski | Charles Mingus | Charles Ives | Charles Bronson | Charles Babbage | Charles III of Spain | Charles Cornwallis, 1st Marquess Cornwallis | Charles Baudelaire | Charles Sanders Peirce |