The eastern portal in Greywell village is the largest winter bat roost in the country, and the second largest colony of Natterer's bat in Europe.
The footpath passes through the towns of New Alresford, Hartley Wintney, Sandhurst, Bracknell and Ascot and the villages of Martyr Worthy, Itchen Abbas, Abbotstone, Upper Wield, Ellisfield, Bradley, Greywell, North Warnborough and Odiham.