They had two daughters; Anna Dorothy (1892-1972), who married Sir Ronald Forbes Adam, 2nd Baronet, and Isabel Mary (1891-?), who married the surgeon Sidney Maynard Smith - her son was the biologist John Maynard Smith.
The importance of behavioral separation as a mechanism for promoting sympatric speciation in a heterogeneous or patchwork landscape is highlighted in John Maynard Smith's seminal paper on sympatric speciation.
He then became a student of John Maynard Smith, still working mostly on the theoretical aspects of evolution and ecology.
They had a daughter Isabel Valentine Maynard Smith (born 1919), and a son, the biologist John Maynard Smith (1920-2004).
Julian Maynard Smith (son of the evolutionary geneticist John Maynard Smith) and Miranda Payne founded the company in 1980, and Maynard Smith remains its artistic director.
Later in the discussion this principle is applied to the thickness required of the bones of a large animal, possibly the first quantitative result in biology, anticipating J. B. S. Haldane's seminal work On Being the Right Size, and other essays, edited by John Maynard Smith.
Collins studied biology at Sussex University, where his tutor was the leading theoretical biologist John Maynard Smith.
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Evolutionarily stable strategies were defined and introduced by John Maynard Smith and George R. Price in a 1973 Nature paper.
Besides Richard Dawkins and George C. Williams, other biologists and philosophers have expanded and refined the selfish-gene theory, such as John Maynard Smith, George R. Price, Robert Trivers, David Haig, Helena Cronin, David Hull, Philip Kitcher, and Daniel C. Dennett.