He worked with Charles Pérez and Maurice Caullery (1925) at Station biologique de Roscoff, at Collège de France under Emmanuel Fauré-Fremiet (1926), at the "Service de biophysique" of the Institut de biologie physico-chimique of Paris under René Wurmser (1927) and became famous for his joint research with P. Trpinac (1938).
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