After close examination and gene sequence studies in 2010, it was determined that there were actually three different species in the Bolivian area: M. brevicaudata and M.domestica.
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Small (less than thirty individuals) colonies were known in Manor and Silver lakes in Bucks county, the Tinicum wetlands in Philadelphia and Delaware counties, the West Branch of Conococheague Creek in Franklin County and possibly Springton Reservoir in Delaware county (McCoy 1985).