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In the 1870s, with Wilhelm Ebstein (1836–1912), he performed important research involving the physiochemical behavior of pepsin in the digestive tract.
Prior to receiving his doctorate, he worked during 1920-21 at Jefferson Medical College as an assistant in physiological chemistry and then worked during 1922-6 as a biochemist at Philadelphia General Hospital (both in Pennsylvania).
Christian de Duve, then chairman of the Laboratory of Physiological Chemistry at the Catholic University of Louvain in Belgium, had been studying the mechanism of action of a pancreatic hormone insulin in liver cells.
His other two sons were also successful scientists, with the pharmacology and physiological chemistry professor Erich Harnack (1853–1914) and the history of literature professor Otto Harnack (1857–1914), father of Arvid Harnack and Falk Harnack.
He worked in the lab of Peter Heitmann, and his father, Samuel Mitja Rapoport, was head of the Institute of Physiological Chemistry.