The early period was dominated by demographic studies such as the work of Benjamin Gompertz and Pierre François Verhulst in the early 19th century, who refined and adjusted the Malthusian demographic model.
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This species, unlike softer shelled physid snails, grows very slowly, and has the lowest intrinsic rate of increase (this means that populations grow very slowly), along with Elimia virginica, in this environment.
He is also interested in life history and population dynamics of exploited or threatened reptile taxa, especially among Varanidae and Crocodilia.
After graduation, he headed to the University of Oxford to study insect population dynamics at the Bureau of Animal Population under Charles Sutherland Elton.