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In mesic, grassland habitats (e.g. Kwa-Zulu Natal Midlands; Wirminghaus & Perrin, 1993; Pretoria Highveld; Brooks, 1974; Zimbabwe grassland; Choate, 1972) and semi-succulent thorny scrub (e.g. Eastern Cape; Perrin, 1980a, b) animals are solitary, with females rearing their litters on their own, and both sexes maintain territories that overlap the territories of the opposite, but not the same, sex (Schradin & Pillay, 2005a).