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The book examined the role of the cell surface in embryology, development and evolution, and presented a critique of gene theory, particularly the views of Jacques Loeb.
Besides Richard Dawkins and George C. Williams, other biologists and philosophers have expanded and refined the selfish-gene theory, such as John Maynard Smith, George R. Price, Robert Trivers, David Haig, Helena Cronin, David Hull, Philip Kitcher, and Daniel C. Dennett.