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Gulella systemanaturae was found in leaf litter in Syzygium forest in the Dedza Mountain in altitudes approximately from 1700 m to 2100 m above sea level.
First described in 2013, two specimens of Pheidole loki were collected in forest leaf litter in 470 and 630 m elevation on Mt. Combani and Mt. Benara, Mayotte, where it occurs in sympatry with P. jonas.
This nocturnal species spends its days in burrows, under logs, or hidden in leaf litter.
They can be found almost everywhere, particularly on flowers, foliage, and in leaf litter, but are often overlooked due to their very small size, which includes the smallest of all known insects (Dicopomorpha echmepterygis).
This provides an excellent camouflage on mulga trees, its usual habitat, it is also found beneath mulga leaf litter.
In a single leaf-litter sample collected in São Bonifácio, Santa Catarina, six workers and two ergatoid gynes very similar to the workers, were captured, which suggests that S. stali is polygynous, as already described for other Sphinctomyrmex species.
The Shovel-billed Kookaburra digs through leaf litter for worms and other prey, and the Vanuatu Kingfisher feeds exclusively on insects and spiders.