It is pumped across the hepatic canalicular membrane into the bile by the transporter MRP2.
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His areas of expertise includes a major contribution to the discovery of P-glycoprotein (MDR1, ABCB1), the multidrug resistance efflux transporter associated with clinical resistance to anti-cancer agents.
He is best known for his discovery of P-glycoprotein, one of the proteins responsible for multidrug resistance.