427 Squadron started as a bomber squadron formed at Croft, England on 7 November 1942 and spent its wartime entirely in England as a part of No. 6 Group RCAF, RAF Bomber Command.
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427 flew Vickers Wellington Mk IIIs and Mk Xs from its first operational mission on 14 December 1942, a minelaying sortie to the Frisian Islands, until May 1943 when it was relocated to Leeming, North Yorkshire.
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