He performed, with the most modern techniques of the time, the first experimental analysis of the action of digitalis on warm-blooded animals and explained its effects, thereby increasing the drug's therapeutic use and value.
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Aroused by interests of physiological drug interaction, he traveled to the European continent and spent a year of associated study under Oswald Schmiedeberg at Strassburg, Austria and six months in Berne under Hugo Kronecker, from whom he learned elements of physiological technique.
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He was also interested for several years in the physiological action of optical isomers and, in c. 1900, the mechanisms of kidney secretion, providing three advanced papers on the subject between 1901 and 1904 in the Journal of Physiology.
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