Root nodules that occur on non-legume genera like Parasponia in association with Rhizobium bacteria, and those that arise from symbiotic interactions with Actinobacteria Frankia in some plant genera such as Alnus, vary significantly from those formed in the legume-rhizobia symbiosis.
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An interaction with a novel RNA-binding protein MtRBP1 (Medicago truncatula RNA-binding protein 1) investigated in the development of Root nodule suggests ENOD40 has a function of cytoplasmic relocalization of nuclear proteins.