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He is credited with the discovery of sexual organ development in Chironomus which eventually led to the general theory on the autonomy of the germ cell.
Between 1850 and 1855, Robert Remak had further refined the germ cell layer concept, and introduced into English were the terms "mesoderm" by Huxley in 1871 and "ectoderm" and "endoderm" by Lankester in 1873.
Spermatocytic seminomas are not considered a subtype of seminoma and unlike other germ cell tumours do not arise from intratubular germ cell neoplasia.